Personally, I doubt it is controlled by a single factor (e.g., diet, flow, etc.). FWIW, I participated in some anemonefish habitat preference research in the northern Red Sea a year and a half ago and noticed some interesting bulb-tip-related patterns. On the reefs, the BTAs that hosted only a single anemonefish, or only juvenile anemonefish, displayed the bulb-tip morphology. On the contrary, the BTAs that hosted an adult pair usually displayed the long skinny morphology. Furthermore, when we removed the fish pair from the BTA, it immediately contracted a bit and quickly developed the bulb-tips. Upon reintroduction of the fish several days later, the BTA re-expanded and eventually lost its bulbs. It's a very interesting phenomenon!
-J