The reef is really a multiple factorial complex system that has many organisms. Each with its own changing needs in food , water parameters , space , oxygen , light. , water flow , Etc
We only measure alkalinity, calcium. , salinity , magnesium , pH. , Nitrate. Phosphate. , and maybe ammonia and nitrite
We leave out the other 20 to 50 trace elements , other perhaps unknown but crucial parameters that may cause an imbalance or livestock death
But overall. As long as we get close to the big parameters. Then we are good to go and able to keep many species alive
Still there are challenges ahead
No one measures the laundry list of crucial amino acids , elements , organic compounds , etc. That each fish and each coral and each snail or crab or clam needs. No one knows. We can make educated guesses and set hypothesis. But the myriad of complex interactions is just too complex
For example : us humans have a pain pathway that involves at least 9 steps “a cascade “ . Each step requires a factor and other things to make the next step happen.
A simple lower animal may not need as many steps or compounds to have said effect. But multiply this by so many complex animals and interactions. Then we get an inkling that we are just understanding maybe 10 or 20 % of what goes on inside corals and fish and their inter species and intra species interactions.