Ken Feldman & Sanjay Joshi research paper has detailed a study of bacteria in a reef tank. I have read Feldman’s initial research, but this is a differrent paper with Sanjay Joshi. It might open some eyes on nutrient management in a reef tank.
Bacterial Counts in Reef Aquarium Water: Baseline Values and Modulation by Carbon Dosing, Protein Skimming, and Granular Activated Carbon Filtration - What are the bacteria populations in the water column of reef tanks, and how does that value compare with bacterial counts in authentic reef...
reefs.com
[Finally, mechanical filtration in the form of skimming but not GAC does provide an effective means of bacteria export, at least up to a point. It appears likely that some types of bacteria are indeed “skimmable”, but others are not. Thus, skimming inadvertently provides severe (?) evolutionary pressure to skew the tank’s resident water column bacteria population to favor the “non-skimmable” cohort.]