Ok I'm going to just brain dump on you here for a minute or two and if something doesn't make sense or you need clarification just ask.
Much visible Alage, Chaeto doesn't seem to be out competing tank.
BTA tend to walk when they get mad, yours are staying put and shrinking. This is abnormal, for this particular species, but not for most other kinds of nems.
Darking Zoos is to be expected with proper lighting, which would explain why they aren't moving. That said your MAX photo period may be lasting too long as in you might need a higher ramp and a shorter max, ie closer to 4 hours.
It seems odd that ALL 3 stopped eating at the exact same time after eating successfully for a few weeks. This leads me to think something happened in the things that are common to them all ie the food source or in the tank. Something sprayed into the house that drifted into the tank...etc. Maybe a bag of Carbon for a week could help here.
Something may have made them sick and it is spreading, but would expect much drooping and mad nems or at a minimum walking because of BTA.
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So yeah, the darkening tells me your light strength is not a bad thing, the odd thing here is typically if the photo period was too long they would just get up and move which yours aren't doing for some reason. I would definitely shorten the photo period and think back and look around and see if something in the tank is off or changed. Be it flow pattern...etc.