Corals exposed to the imbalanced, HN/LP conditions, displayed a smaller polyp size and a bleached appearance that closely resembled the phenotype observed in low nutrient water (LN/LP) (Figs. 1, 2a). In contrast, the corals kept under LN/HP imbalanced nutrient levels showed a similar phenotype to the nutrient replete (HN/HP) treatment. The bleached appearance of the polyps from HN/LP conditions was associated with low numbers of zooxanthellae in the tentacle tissue (Figs. 1, 2b, Table 1), similar to the low nutrient LN/LP treatment. In contrast, the symbiont numbers in the tissue of LN/HP exposed corals were comparable to those of corals from nutrient replete (HN/HP) conditions